Last modified on 03 September 2002.


                 ITS1 -->
Heliantheae    ||UCGAA-+CCU--GCA+A--G-C-AG-AA-C+AC++GUGAACA+GU+A-
Tageteae       ||UCGAA-UCCU--GC++A--G-C-AG-AA-++AC+UGUGAACA+GUAA-
Eupatorieae    ||UCGAA-+CCU--G++++--G-C-A+-AA-CAAC++GUGAAC+UGUA+-
Helenieae      ||UCGAA-+CCU--GCA+A--G-C-AG-AA-C+ACC+GUGAACA+GUAC-
Astereae       ||UCGAA-GCCU--GCAAA--G-C-AG-AA-CGACCCG+GAACAUGUUA-
Gnaphalieae    ||UCGAA-CCCU--GCAAA--G-C-AG-AA-CGACCCGUGAACA+GUAA-
Inuleae        ||UCGAA-GCCU--AUAAA--G-C-A+-AA-+GACCCGUGAACGUGUAA-
Senecioneae    ||UC+AA-ACCU--GCAU+--G-C-++-AA-++AC++GUGAACAUGUAA-
Anthemideae    ||UCGAA-CCCU--GCAAA--G-C-AG-AA-CGACCCGUGAACACGUAA-
Cardueae       ||UCGAA-GCCU--GCACA--G-C-AG-AA-CGACCCGUGAACA+GUAA-
Vernonieae     ||UCGA+-++++--GC+++--U-+-++-AA-++AC++GUGAACA+GU++-
Lactuceae      ||UCGAA-CCCU--GCAA+--G-C-AG-A++CGACC+G+GAAC+UGUA+-
Mutisieae      ||UCGAA-+CCU--GC++A--G-+-AG-AA-CGACC+G+GAAC+UGUA+-
Barnades.      ||U+GA+-ACCU--GCA+A--G-+-AG-AA+CGACC+G+G+ACA+GUA+-

                                                 <-- Helix 1A -->
Heliantheae    ++A---CA+C+--GGC-+U+-+++--+GG-++-++A-AGC---+U+U-G+
Tageteae       +AA---CA+CA-U+GC-U+U-+++--+G+-+U-+G+-UGC---UU+U-G+
Eupatorieae    CAA---CAA++-UGGC-+U+-+CG--GG+-++-+GA-++C--+U++U-GU
Helenieae      +AA---CAAC+-UGGC-+U+-+++--+GG-A+-++A-+++---+UUU-++
Astereae       +AA---CAACC-AUGC-CA+-GA---UG+-GU-CG+--GC---+U++-GU
Gnaphalieae    +UA---C+A+C-GGGC-AAC-+UA--+GG-AU-UGA--GC---UU+U-G+
Inuleae        CAA---CA+C+-++++-+UC-AUA--G+G-AU-C+G--+C---UUUU-G+
Senecioneae    ++A---+A+U+-+GGU-GU+-CU+--GG+-AU-++G--GC---++U+-+U
Anthemideae    +AA---CAAC+-GAG+-GUC-GAG--UGG-+U-++A--GC---+++U-GU
Cardueae       UCA---CAA++-GGG+-GUC-G++--GGG-AU-+GG--GU---G+GA-GC
Vernonieae     U++---+AA+C-++G+-+++-GG+--+++-A--+GG--GC---U+++-GC
Lactuceae      ++A---+AA++-+GG+-G++-+++--G++-A+-++G--+C---+UU+-G+
Mutisieae      ++A---+++++-GGG+-GU+-G++--GG+-AU-CGG--++---++++-++
Barnades.      +A+-++CA+CA-+G++-G+C-+CG--+++-++-++G--G+---++++-+C


Heliantheae    U-U+G-++----CCU+-GU-+A-++--CCU+--GU+G+-C+-UG-+GU-U
Tageteae       +-U++-A+----CCU+-GU-++-++--C++U--GUC+A-++-UG-UGU-U
Eupatorieae    +-UC+-A+----CC++-G+-++-+G--+C+U--GU++A-CG-UG-+G+-U
Helenieae      ++U+G-++----+CU+-G+-G+-+G--CC++--GUCG+-C+-UG-UGU-+
Astereae       --UCG-AU----C+UC-+U-GG-CA--CACC--GU+GA-UG-UG-C++-C
Gnaphalieae    --UUG-AU----CCUU-AG-UU-+C--CUUU--GUC+A-UG-UG-+GU-U
Inuleae        --U++-+G----ACCU-AU-G+-C+--CCUC--GUCGA-UU-UG-CAU-C
Senecioneae    --CC+-AU----UC+U-+G-GA-+G--CCA+--GUU+A-+G-UG-CGU-C
Anthemideae    --UUG-A+----CCUC-U+-GA-+G--CUUU--GUCGA-UG-+G-CAU-U
Cardueae       --C++-AG----+C+U-+C-GA-UG--CU++--G++GG-CA-UG-CGU-+
Vernonieae     --+++-++----++++-++-++-++--CC+U--GC++A-C+-+A-C++-U
Lactuceae      --+++-++----+U++-++-++-++--CC++--++CGG-++-U+-++U-+
Mutisieae      --++G-+G----A+C+-++-G+-+G--C+C+--G+CG+-CG-UG-C++-+
Barnades.      --C++-++----A++C-GU-+U-C+-++C++--++CG+-UG-+G-+G+-+

               <--                 Helix 1B                   -->
Heliantheae    C-AUG+--+++++-+++++C+++-++G+G++----UC+UG-GAU+-UCA+
Tageteae       C-AUG+--------+++++++++-+++G+++----+++UG-GACA-UUA+
Eupatorieae    U-++GG-------++++++++UU-+++++++----+++++-++CA-UCA+
Helenieae      +-+UG+--------++++C+U++-++GG+++----UCAUG-G++A-U++U
Astereae       C-U+G+--------+++++++++-+++G+++----++UUG-GU+G-U+GC
Gnaphalieae    +-GAGA------------CUCCU-UAG+GA+----GA++G-+ACG-UCAC
Inuleae        +-AUGG--------++GCCUCUU-CGGAGC+----UCC++-G+UG-UCAA
Senecioneae    U-UUGG-------++++++++++-U++++U+----C+A++-GA+G-U+A+
Anthemideae    +-AC++--------+++GU+CU+-UUGG+C+----U+GUG-A+UG-U+UC
Cardueae       C-A+GG--------UGCCU+++U-++AGGCA----UC+UG-GA+G-UU+U
Vernonieae     U-+U++------+++++++++++-+++++++----+++++-++UG-U+U+
Lactuceae      +-+UGG--------U++C+++UU-+++++U+----+CAUG-GAU+-U+++
Mutisieae      +-+U++-------+++++C+U+U-++GGG++----++++G-+++G-U+U+
Barnades.      +-++++-------++++C+++++-+++G+++----+++++-++++-++G+

                                             <-- Helix 1C -->
Heliantheae    -GU+-GACA-+A++-+AA-CAA++++CC--GGCAC+A+A+GUG-CC-A-A
Tageteae       -GUU-GGCA-+AA--CAA-C+++-CCCC--GGCACAACACGUG-CC-A-A
Eupatorieae    -G+U-GACG-CAA--CAA-C+A+-CCCC--GGCACA+CACGUG-CC-A-A
Helenieae      -GU+-GGCA-CA+--+AA-CAA++CCCC--GGCACGGCA+GUG-CC-A-A
Astereae       -+UU-GACG-UAA--CAA--AA--CCC+--GGCACGGGAUGUG-CC-A-A
Gnaphalieae    -AU+-GGCA-+AC--UAA-CCAA-CCCC--GGCACGGAAUGUG-CC-A-A
Inuleae        -AAA-GGC+-+AA--CAA-++AA-+CCC--GGCACGGCAUGUG-CC-A-A
Senecioneae    -+U+-G+C+-+AA--C++-CAA++CCCC--GGCAC+++AUGUG-+C-A-A
Anthemideae    -+U+-GGCG-CAA--UAA-CAAC-CCCC--GGCACAAUG+GUG-CC-A-A
Cardueae       -GUC-GGCA-C+A--AAA-CAAA+CCCC--GGCACG+CAUGUG--C-A-A
Vernonieae     -GU+-+GCA-+++--AAA-CAAA+CCCC--GGCACGGAA+GUG-CC-A-A
Lactuceae      -+++-GG+A-+++--UAA-CAAA-CCCC--GGCACG++AUGUG-CC-A-A
Mutisieae      -GUC-G+C+-+++--UAA-CAAA-CCCC--GGCACGG+A+GUG-CC-A-A
Barnades.      -++C-G+C+++C+--UAA-C+AA-CCCC--GGCACGG+++GUG-CC-A-A


Heliantheae    GGA--AAA-C+++-ACUU---AAAGG--GC++GU--GC-++U++C-GCCC
Tageteae       GGA--AAA-+UA+-A+++--++AAG+--GC+++U--GC-++UGA+-GCCC
Eupatorieae    GGA--AAA-C+AA-ACUU--AAGAG+--G+++GU--GC-++UGA+-+CCC
Helenieae      GGA--AAA-C++A-ACUU--AAGA+G--G+++GU--GC+++UGAC-+CCC
Astereae       GGA--A++-+UAA-AUU+--AAGAAU+-GC++GU--+C-CAUGA+-GUCC
Gnaphalieae    GGA--AAU-++UA-ACUU-AAAGAAU+-GA+GGU--UU-CAUGAU-+UCC
Inuleae        GGA--AAA-+++A-ACUU-UA+GAAGG-C+UCAU--G+-CAUG+U-+CCC
Senecioneae    GGA--AA+-AUAA-A++U--+++AAGG-GCUUGU--++-++++C+-++++
Anthemideae    GGA--AAA-CUAA-ACU+--+AGAAG--GCU+GU--UU-CAUGUU+GCCC
Cardueae       GGA--AAA-C+AA-AC++--AAGAAGG-G+GCGU--C+-CGUGUU-GCCC
Vernonieae     GGA--AAA-+AAA-A++U--AAGAAGG-+U++G+--C+-+GUGA+-G+++
Lactuceae      GGA--AAA-++A+-A+A+--+AGAAGG-++++++--C+-++++U+-GCCC
Mutisieae      GGA--A++-+++A-AC+U--AAGAA+A-GC+CGU--C+-+G+GU+-+CCC
Barnades.      GGA++A++++++++ACU+--+A+AAGA-GCACG+--++-C+++++-G++C


Heliantheae    C---GUU+++CGGU+-+GCG--+++U-G+++-GU-GGC+UCUUU--GU-A
Tageteae       C---GUU-U++GGUG-UGC+--CAUU-GUAU-GU-GGCUUCUUU--++-A
Eupatorieae    +---GUA-+++GGUG-+G+U--C+U+-GUA+-G+-GGCUUCUUU--GU-A
Helenieae      +---GUU-U+UGGUG-UG++--CAUU-GU++-GU-GGCUUCUUU--+U-A
Astereae       C---GUU-+GCGGUG-UGCU--CAUG-G+G+-GU-GGC+UCUUU--+U-A
Gnaphalieae    C---GUUUU+CGGUG-++CU--CAUG-AAAC-U+-UAC+UCUUU--GU-A
Inuleae        C---GUU-CGCGGUG-UGUG--CAUG-GGA+-G+-GGCUUCUUU--+U-A
Senecioneae    +---GUU-++CGGUG-AUUG--++++-++AU-CU-+GCUUCUUU--A+-A
Anthemideae    C---GUU-CGCGGUG-UGCU--CAUG-G+A+-GU-GGCUUCUUU--AU-A
Cardueae       C---GUU-++CGGUG-UGCA--CA+G-GGUC-GU-GGCCUCUCA--UU-A
Vernonieae     +++-+UU-+++++++-++++--++++-G+++-+U-++++++UU+--++-A
Lactuceae      C---GUU+++CGGUG-UGC+--U+++-G+++-GU-+GCCU+CUU--++-A
Mutisieae      C---G+U-+GCGGUG-UGC+--+G+G-G++++GU-+GCUUCUU---A+-A
Barnades.      C---GUU-+GCGGUG-++++--+++G-+++C+G+-+GCUUCUU+--++-A

                       5.8S -->
Heliantheae    A-+C-+||++++++++++++++++++++++++++++++++++++++++++
Tageteae       A-UC-A||+AAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU
Eupatorieae    A-UC-+||U+++++++++++++++++++++++++++++++++++++++++
Helenieae      A-UC-+||UAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU
Astereae       A-UC-A||CAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU
Gnaphalieae    A-UC-A||CAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU
Inuleae        A-CC-A||++++++++++++++++++++++++++++++++++++++++++
Senecioneae    A-U+-A||++++++++++++++++++++++++++++++++++++++++++
Anthemideae    A-++-+||+++++++++++++++++++++++++++++++++---------
Cardueae       A-CC-A||+A+++++++---------------------------------
Vernonieae     A-++-A||++++++++++++++++++++++++++++++++++++++++++
Lactuceae      A-++-A||C+++++++++++++++++++++++++++++++++++++++++
Mutisieae      A-UC-A||++++++++++++++++++++++++++++++++++++++++++
Barnades.      A-+C-A||CAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU


Heliantheae    ++++++++++++++++++++++++++++++++++++++++++++++++++
Tageteae       GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA
Eupatorieae    ++++++++++++++++++++++++++++++++++++++++++++++++++
Helenieae      GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA
Astereae       GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA
Gnaphalieae    GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA
Inuleae        ++++++++++++++++++++++++++++++++++++++++++++++++++
Senecioneae    ++++++++++++-+++++++++++++++++++++++++++++++++++++
Anthemideae    -----------------+++++++++++++++++++++++++++++++++
Cardueae       --------------------------------------------------
Vernonieae     ++++++++++++++++++++++++++++++++++++++++++++++++++
Lactuceae      ++++++++++++++++++++++++++++++++++++++++++++++++++
Mutisieae      ++++++++++++++++++++++++++++++++++++++++++++++++++
Barnades.      GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA


Heliantheae    ++++++++++++++++++++++++++++++++++++++++++++++++++
Tageteae       ACCAUCGAGUUUUUGAACGCAAGUUGCGCCCGAAGCCAU++GGUUGAGGG
Eupatorieae    ++++++++++++++++++++++++++++++++++++++++++++++++++
Helenieae      ACCAUCGAGUUUUUGAACGCAAGUUGCGCC+GAAGCC+UCUGG++GAGGG
Astereae       ACCAUCGAGUUUUUGAACGCAAGUUGCGCCCGAAGCCAUUCGGC+GAGGG
Gnaphalieae    ACCAUCGAGUUUUUGAACGCAAGUUGCGCC+GAAGCCAUUUGGUUGAGGG
Inuleae        ++++++++++++++++++++++++++++++++++++++++++++++++++
Senecioneae    ++++++++++++++++++++++++++++++++++++++++++++++++++
Anthemideae    +++++++++++++++++++++++++++++++++++++++++++++++-++
Cardueae       --------------------------------------------------
Vernonieae     ++++++++++++++++++++++++++++++++++++++++++++++++++
Lactuceae      ++++++++++++++++++++++++++++++++++++++++++++++++++
Mutisieae      ++++++++++++++++++++++++++++++++++++++++++++++++++
Barnades.      ACCAUCGAGUUUUUGAACGCAAGUUGCGCCC+AAGCCA++CGGC+GAGGG

                                       ITS2 --> <-- Helix 2A
Heliantheae    ++++++++++++++++++++++||+-UCA+G--UCGC-CCC---+A-CCA
Tageteae       CACGUCUGCCUGGGCGUCACGC||A-UC+UG--U++C-CC+---+A-CCA
Eupatorieae    ++++++++++++++++++++++||A-UCA+G--U+GC-CCA---CA-+CA
Helenieae      CACGUCUGCCUGGGCGUCACGC||A-UCACG--UCGC-CC+---CA-C+A
Astereae       CACGUCUGCCUGGGCGUCACGC||A-UCGCG--UCGC-UCC---C+-C+A
Gnaphalieae    CACGUCUGCCUGGGCGUCAC+C||A-UCGUG--UCGC-CCC+--+A-CCA
Inuleae        ++++++++++++++++++++++||+-+++++--UCGC-UCCU--C+-+C+
Senecioneae    ++++++++++++++++++++++||+-U+G+G--UC+C-C++---C+-+++
Anthemideae    ++++++++++++++++++++++||A-UCGC+--UCGC-CCCC--++-CAA
Cardueae       ----------------------||A-UCGCG--UCGC-CCC+--+A-+CA
Vernonieae     ++++++++++++++++++++++||A-U+GC+--UCGC-C+C+--+U-+++
Lactuceae      ++++++++++++++++++++++||A-UC+C+--UCGC-CCC+--++-+CA
Mutisieae      ++++++++++++++++++++++||A-UCGC+--UCGC-CCCC--+A-CAA
Barnades.      CACG+CUGCCUGGGCGUCACGC||A-UCGCG--UCG+-CCC++-+A-CAA

                                 Helix 2A                     -->
Heliantheae    +--+++U----CC+---+U+++++G+++U+-----UU+++U-+GGG-GCG
Tageteae       +--+++U----CC+---++++++GG+U+U+-----+UG+++-+GGG-G+G
Eupatorieae    +--ACAU----CC+---+++++++++++UG-----+UGUAU-GUGG-GCG
Helenieae      +--+C+U----CC+---++++++GGAU+++-----+U+G+U-G+GG-GCG
Astereae       +--++++----+C+---UU--++GG-AUG+-----UUGG+U-GGG+-GCG
Gnaphalieae    +--+++U----C++---UUA-+++GAUGUU-----U+GUG+-+GGG-GCG
Inuleae        U--GC+U----CCU---+-AAA+GGGUG+G-----C++GAU-A+GA-GCG
Senecioneae    +-++C++--++++U---U+A+GGGG+UGU+-----U++++U-GGGG+G+G
Anthemideae    +--U+++-----++---+++++++++++++-----U+U+U+-GGGG-GCG
Cardueae       +--++++----C++---++++G++++++GU-----+U+GU+-UGGG-+CG
Vernonieae     +--GC+U--+++++---A+++++A++C++------+U++UU-GGGG-GCG
Lactuceae      +--+C+U+---C++---+++++G+++++++-----+G+++U-+GGG-GCG
Mutisieae      +--G++U+---CC+---++++GG+++++++++---+++G++-GGGG-GCG
Barnades.      C--+CA++---C++---+++++GCG++++------++++++-+GGG-GCG

                        <--            Helix 2B               -->
Heliantheae    G-A+--AU-U-GGUC-UC-C+G-UGCC-+AU---GG+G-+GG-UU-GGCC
Tageteae       +-A+--AU-U-GGUC-UC-C+G-UGCC-+++++-GG+G-++G-UU-GGCC
Eupatorieae    G-AG--AC-U-GGUC-UC-CUG-UGCC-+A+---GGUG-CGG-UU-GGCC
Helenieae      G-AG--AU-U-GGUC-UC-C+G-UG++-++U---G+++-+GG-UU-GGCC
Astereae       G-AU--A+-U-GGUC-UC-CCG-UUU+UCAC---CGAG-CGG-UU-GGCC
Gnaphalieae    G-AG--AU-U-GGUC-UC-CCG-+UUC-U++---GAUA-CGG-UU-GGCC
Inuleae        G-AU--A+-U-GGUC-UC-CCG-UGCC-UA+---GGUG-+GG-UU-GGC+
Senecioneae    G-AG--AU-U-GGU+-UC-CCG-UU+C-U++---G+UG-CGG-UU-GGCU
Anthemideae    G-AU--AU-U-GGUC-UC-CCG-UGCU-CA+---GG+G-UGG-UU-GGCC
Cardueae       G-A+--A+-U-GGUC-UC-CCG-UG++-+++---GG+G-+GG-UU-GGCC
Vernonieae     G-A+--AU-U-GG+C-UC-CCA-UGC+-+A+---GG+G-UGG-UU-GGCC
Lactuceae      G-A+--AU-U-GG+C-UC-CCG-U+C+-U++---+GU+-+GG-UU-GGCC
Mutisieae      G-A+--A+-U-GG+C-U+-CCG-UGC+-++++--GG+G-++G-UU-GGCC
Barnades.      G-A+--++-U-GG+C-UC-CCG-UG+C-+++++-+G+G-+GG-+U-GGCC

                                               <--   Helix 2C
Heliantheae    UA-AA-UA+-+AGUC+CCU-C++-GA------G+-GAC-GCACG+-C-U-
Tageteae       UA-AA++A+-GAGUC++C+-+AA+++------++-GA+-GCACGA-C-U-
Eupatorieae    CA-AA-+A+-GAG+CC++U-UAA-GA++----G+-GAC-GCACGA-C-U-
Helenieae      UA-AA-UA+-GAGUC+++U-C+A-GA------+G-GA+-GCACGA-C-U-
Astereae       +A-AA-+AA-+AG++CC++-UU--GA------CG-GG+-+CA+GA-C-U-
Gnaphalieae    +A-AA-UAG-GAGUCCCCA-UC--GA------UG-GAC-GCACGA-C-U-
Inuleae        AA-AA-UAG-+AGUCU+CU-U+--GA------+G-GAC-ACACG+-C-A-
Senecioneae    +A-A+-U++-GA+U+C++U-U++++A------+G-+AC-+C+CGA-U-U-
Anthemideae    +A-AA-UA+-GAGU-CCCU-UC--GA------UG-GAC-GCAC+AAC-U-
Cardueae       UA-AA-AA+-GAGUCCCCU-U+--G+------+G-G++-GCACGG-C-U-
Vernonieae     +A-AA-++G-AAGUCCU++-UC--G+------U+-G+U-GCA+GA-C-U-
Lactuceae      +A-AA-+++-GAGU++CCU-++--GG------+G-GA+-GCAC+A-C-U-
Mutisieae      UA-AA-+++-GAGUCC++U-U+--G+------+G-GAC-GCAC++-C-U-
Barnades.      +A-AA++++-G+GUC+C++-++--GA------++-+++-+CAC++-C-+-

                                Helix 2C                      -->
Heliantheae    ---AGUGGUGGUU-GAUAA-GAC+---G-UCGU+UC--G+GU-+GUG+GU
Tageteae       ---AGUGGUGGUU-UGUAA-+A+A---G-UCGUCUC--GUGU-UGUG++U
Eupatorieae    ---GGUGGUGGUU-GAUUA-+AC+---G-UCGUCUC--GUGU-CGUG+G+
Helenieae      ---AGUGGUGGUU-GAU+A-+ACA---G-UCGUCUC--GUGU-CGUG+GU
Astereae       ---A+UG+UGGU+-GACAA-AACC-----++GAAU+--++GU-+G+GUGU
Gnaphalieae    ---AGUGGUGGUU-GACAA-AACC---U-UCGUCUU--G+GU-+GUGCGU
Inuleae        ---AGUGGUGGUU-GA+AA-AACC---U++AGUCUC--GUGU-+GUGUGU
Senecioneae    ---AGUGGUGGUU-G++AA-GACC---+-UCU+AU+--GAGU-+GUG+GU
Anthemideae    ---AGUGGUGGUC-GUAAA-AACC---C-UCGU++U--UUGU-U++G+G+
Cardueae       ---AGUGGUGGUU-GU+AA-GGCC---U-UCGUAUC--GAGC-CGUG+--
Vernonieae     ---A+UGGUGGUU-+++AA-GACC---U-++G+U++--++GU-+++GU++
Lactuceae      ---AGUGGUGGUU-GAA+A-GACC---C-UCGUCUU--+UGU-+GUG+GU
Mutisieae      ---AGUGGUGGUU-G+CAA+G+CC---+-U+++UUC--+UG+-CGUGCGU
Barnades.      ---AGUGGUGG++-+++AA-+GCC-----++AGUUC--GUGU-CGUG++U


Heliantheae    U-+---+++--+++U+++G++++++++CU+--+U+++-++A--CC+-+++
Tageteae       +-+---U+A+-+++++++++G++A++++U+--U+A++-+++++CCC--++
Eupatorieae    +-U---+GA--UUCUUG-A++G+A+A+++++-UU++A-+UA--CCC-U+A
Helenieae      U-+---++A--+UCUUGA+G+++A++ACUC--UUAAA-++A--CCC-+G+
Astereae       C-U---+G---UCAAAA-GG+UGC+UCUUA--++----+GA--CCC--AA
Gnaphalieae    +-+---U+AG-UCGUA+-GGGAAGAU-CUC--UUUAA-AGA--CCC-CAA
Inuleae        C-C---+GA+-UUGUAA-GCGAAGA-CC+++-UA++--+UA--CCC-UAA
Senecioneae    U-C---++A+-++G+AA--+G+A+A+-C+C--U++AA-U+A--+CC-U+A
Anthemideae    +-+---+++G-UCG+AA-GG+AA++--CUC--U+++A-A+A--CCC-+AA
Cardueae       ------++A+-++G++A-GGGA+++G-CUC--U++AA-AGA--+CC-+AA
Vernonieae     C-+---++AG-+CG+AA-G++A+A++-+UU+-+++AA-AG+--CC+-U++
Lactuceae      +-G---U+AG-C+G++A-+GGA+G++-+U+--+++++-AGA--CCC-+A+
Mutisieae      +-+---+++G-UCGC+A-++++++-G++++++G++A--AGA--CCC-+AA
Barnades.      +-+---++++-+CG+++--+GG++++-CUC+-++++--AGA--CCC+++A

                  <--      Helix 2D      -->
Heliantheae    +--G+-GU-UGU-C-+U-++-GA+GA+GC-UUC-GA++||
Tageteae       U--G+-+U-UGU-C-UU-++-GA++AUGC-UUC-GAUC||
Eupatorieae    U--G+-G+-+GU-C-UU-+U-G++++CUC-UU+-GAU+||
Helenieae      +--G+-GU-UGU-C-UU-+U-GAUG++GC-UU+-GA--||
Astereae       +--GC-GU-UGU-+-+U-++-+A+GA+GC-UUC-GA++||
Gnaphalieae    U--G+-GU-UGU-C-UU-U+-GAUGAC+C-UUC-G+++||
Inuleae        C--GU-GU-+GU-C-++-+U-GAC+ACGC-UUC-GACC||
Senecioneae    U--GU-G+-C+U-+-U+-+U-+A+GA+GC-U++-GA++||
Anthemideae    +--GU-GU-+GU-C-UU-++-GA+GACGC-UUC-GA++||
Cardueae       +--GU-GU-+GU-C-UU-++-GACGA+GC-UUC-GA--||
Vernonieae     U--G+-+U-+G+-+-U+-++-++++A+G+-UUC-G+++||
Lactuceae      U--GU-+U-+GU-+-++-++-+A+GAU++-+UC-GAC+||
Mutisieae      +--G+-GU-CGU-C-++-++-GACG++G+-UU+-GACC||
Barnades.      +--++-G+-+++-C-U++++-G++G+++C-U+C-GAC+||