Last modified on 03 September 2002.
ITS1 --> Heliantheae ||UCGAA-+CCU--GCA+A--G-C-AG-AA-C+AC++GUGAACA+GU+A- Tageteae ||UCGAA-UCCU--GC++A--G-C-AG-AA-++AC+UGUGAACA+GUAA- Eupatorieae ||UCGAA-+CCU--G++++--G-C-A+-AA-CAAC++GUGAAC+UGUA+- Helenieae ||UCGAA-+CCU--GCA+A--G-C-AG-AA-C+ACC+GUGAACA+GUAC- Astereae ||UCGAA-GCCU--GCAAA--G-C-AG-AA-CGACCCG+GAACAUGUUA- Gnaphalieae ||UCGAA-CCCU--GCAAA--G-C-AG-AA-CGACCCGUGAACA+GUAA- Inuleae ||UCGAA-GCCU--AUAAA--G-C-A+-AA-+GACCCGUGAACGUGUAA- Senecioneae ||UC+AA-ACCU--GCAU+--G-C-++-AA-++AC++GUGAACAUGUAA- Anthemideae ||UCGAA-CCCU--GCAAA--G-C-AG-AA-CGACCCGUGAACACGUAA- Cardueae ||UCGAA-GCCU--GCACA--G-C-AG-AA-CGACCCGUGAACA+GUAA- Vernonieae ||UCGA+-++++--GC+++--U-+-++-AA-++AC++GUGAACA+GU++- Lactuceae ||UCGAA-CCCU--GCAA+--G-C-AG-A++CGACC+G+GAAC+UGUA+- Mutisieae ||UCGAA-+CCU--GC++A--G-+-AG-AA-CGACC+G+GAAC+UGUA+- Barnades. ||U+GA+-ACCU--GCA+A--G-+-AG-AA+CGACC+G+G+ACA+GUA+- <-- Helix 1A --> Heliantheae ++A---CA+C+--GGC-+U+-+++--+GG-++-++A-AGC---+U+U-G+ Tageteae +AA---CA+CA-U+GC-U+U-+++--+G+-+U-+G+-UGC---UU+U-G+ Eupatorieae CAA---CAA++-UGGC-+U+-+CG--GG+-++-+GA-++C--+U++U-GU Helenieae +AA---CAAC+-UGGC-+U+-+++--+GG-A+-++A-+++---+UUU-++ Astereae +AA---CAACC-AUGC-CA+-GA---UG+-GU-CG+--GC---+U++-GU Gnaphalieae +UA---C+A+C-GGGC-AAC-+UA--+GG-AU-UGA--GC---UU+U-G+ Inuleae CAA---CA+C+-++++-+UC-AUA--G+G-AU-C+G--+C---UUUU-G+ Senecioneae ++A---+A+U+-+GGU-GU+-CU+--GG+-AU-++G--GC---++U+-+U Anthemideae +AA---CAAC+-GAG+-GUC-GAG--UGG-+U-++A--GC---+++U-GU Cardueae UCA---CAA++-GGG+-GUC-G++--GGG-AU-+GG--GU---G+GA-GC Vernonieae U++---+AA+C-++G+-+++-GG+--+++-A--+GG--GC---U+++-GC Lactuceae ++A---+AA++-+GG+-G++-+++--G++-A+-++G--+C---+UU+-G+ Mutisieae ++A---+++++-GGG+-GU+-G++--GG+-AU-CGG--++---++++-++ Barnades. +A+-++CA+CA-+G++-G+C-+CG--+++-++-++G--G+---++++-+C Heliantheae U-U+G-++----CCU+-GU-+A-++--CCU+--GU+G+-C+-UG-+GU-U Tageteae +-U++-A+----CCU+-GU-++-++--C++U--GUC+A-++-UG-UGU-U Eupatorieae +-UC+-A+----CC++-G+-++-+G--+C+U--GU++A-CG-UG-+G+-U Helenieae ++U+G-++----+CU+-G+-G+-+G--CC++--GUCG+-C+-UG-UGU-+ Astereae --UCG-AU----C+UC-+U-GG-CA--CACC--GU+GA-UG-UG-C++-C Gnaphalieae --UUG-AU----CCUU-AG-UU-+C--CUUU--GUC+A-UG-UG-+GU-U Inuleae --U++-+G----ACCU-AU-G+-C+--CCUC--GUCGA-UU-UG-CAU-C Senecioneae --CC+-AU----UC+U-+G-GA-+G--CCA+--GUU+A-+G-UG-CGU-C Anthemideae --UUG-A+----CCUC-U+-GA-+G--CUUU--GUCGA-UG-+G-CAU-U Cardueae --C++-AG----+C+U-+C-GA-UG--CU++--G++GG-CA-UG-CGU-+ Vernonieae --+++-++----++++-++-++-++--CC+U--GC++A-C+-+A-C++-U Lactuceae --+++-++----+U++-++-++-++--CC++--++CGG-++-U+-++U-+ Mutisieae --++G-+G----A+C+-++-G+-+G--C+C+--G+CG+-CG-UG-C++-+ Barnades. --C++-++----A++C-GU-+U-C+-++C++--++CG+-UG-+G-+G+-+ <-- Helix 1B --> Heliantheae C-AUG+--+++++-+++++C+++-++G+G++----UC+UG-GAU+-UCA+ Tageteae C-AUG+--------+++++++++-+++G+++----+++UG-GACA-UUA+ Eupatorieae U-++GG-------++++++++UU-+++++++----+++++-++CA-UCA+ Helenieae +-+UG+--------++++C+U++-++GG+++----UCAUG-G++A-U++U Astereae C-U+G+--------+++++++++-+++G+++----++UUG-GU+G-U+GC Gnaphalieae +-GAGA------------CUCCU-UAG+GA+----GA++G-+ACG-UCAC Inuleae +-AUGG--------++GCCUCUU-CGGAGC+----UCC++-G+UG-UCAA Senecioneae U-UUGG-------++++++++++-U++++U+----C+A++-GA+G-U+A+ Anthemideae +-AC++--------+++GU+CU+-UUGG+C+----U+GUG-A+UG-U+UC Cardueae C-A+GG--------UGCCU+++U-++AGGCA----UC+UG-GA+G-UU+U Vernonieae U-+U++------+++++++++++-+++++++----+++++-++UG-U+U+ Lactuceae +-+UGG--------U++C+++UU-+++++U+----+CAUG-GAU+-U+++ Mutisieae +-+U++-------+++++C+U+U-++GGG++----++++G-+++G-U+U+ Barnades. +-++++-------++++C+++++-+++G+++----+++++-++++-++G+ <-- Helix 1C --> Heliantheae -GU+-GACA-+A++-+AA-CAA++++CC--GGCAC+A+A+GUG-CC-A-A Tageteae -GUU-GGCA-+AA--CAA-C+++-CCCC--GGCACAACACGUG-CC-A-A Eupatorieae -G+U-GACG-CAA--CAA-C+A+-CCCC--GGCACA+CACGUG-CC-A-A Helenieae -GU+-GGCA-CA+--+AA-CAA++CCCC--GGCACGGCA+GUG-CC-A-A Astereae -+UU-GACG-UAA--CAA--AA--CCC+--GGCACGGGAUGUG-CC-A-A Gnaphalieae -AU+-GGCA-+AC--UAA-CCAA-CCCC--GGCACGGAAUGUG-CC-A-A Inuleae -AAA-GGC+-+AA--CAA-++AA-+CCC--GGCACGGCAUGUG-CC-A-A Senecioneae -+U+-G+C+-+AA--C++-CAA++CCCC--GGCAC+++AUGUG-+C-A-A Anthemideae -+U+-GGCG-CAA--UAA-CAAC-CCCC--GGCACAAUG+GUG-CC-A-A Cardueae -GUC-GGCA-C+A--AAA-CAAA+CCCC--GGCACG+CAUGUG--C-A-A Vernonieae -GU+-+GCA-+++--AAA-CAAA+CCCC--GGCACGGAA+GUG-CC-A-A Lactuceae -+++-GG+A-+++--UAA-CAAA-CCCC--GGCACG++AUGUG-CC-A-A Mutisieae -GUC-G+C+-+++--UAA-CAAA-CCCC--GGCACGG+A+GUG-CC-A-A Barnades. -++C-G+C+++C+--UAA-C+AA-CCCC--GGCACGG+++GUG-CC-A-A Heliantheae GGA--AAA-C+++-ACUU---AAAGG--GC++GU--GC-++U++C-GCCC Tageteae GGA--AAA-+UA+-A+++--++AAG+--GC+++U--GC-++UGA+-GCCC Eupatorieae GGA--AAA-C+AA-ACUU--AAGAG+--G+++GU--GC-++UGA+-+CCC Helenieae GGA--AAA-C++A-ACUU--AAGA+G--G+++GU--GC+++UGAC-+CCC Astereae GGA--A++-+UAA-AUU+--AAGAAU+-GC++GU--+C-CAUGA+-GUCC Gnaphalieae GGA--AAU-++UA-ACUU-AAAGAAU+-GA+GGU--UU-CAUGAU-+UCC Inuleae GGA--AAA-+++A-ACUU-UA+GAAGG-C+UCAU--G+-CAUG+U-+CCC Senecioneae GGA--AA+-AUAA-A++U--+++AAGG-GCUUGU--++-++++C+-++++ Anthemideae GGA--AAA-CUAA-ACU+--+AGAAG--GCU+GU--UU-CAUGUU+GCCC Cardueae GGA--AAA-C+AA-AC++--AAGAAGG-G+GCGU--C+-CGUGUU-GCCC Vernonieae GGA--AAA-+AAA-A++U--AAGAAGG-+U++G+--C+-+GUGA+-G+++ Lactuceae GGA--AAA-++A+-A+A+--+AGAAGG-++++++--C+-++++U+-GCCC Mutisieae GGA--A++-+++A-AC+U--AAGAA+A-GC+CGU--C+-+G+GU+-+CCC Barnades. GGA++A++++++++ACU+--+A+AAGA-GCACG+--++-C+++++-G++C Heliantheae C---GUU+++CGGU+-+GCG--+++U-G+++-GU-GGC+UCUUU--GU-A Tageteae C---GUU-U++GGUG-UGC+--CAUU-GUAU-GU-GGCUUCUUU--++-A Eupatorieae +---GUA-+++GGUG-+G+U--C+U+-GUA+-G+-GGCUUCUUU--GU-A Helenieae +---GUU-U+UGGUG-UG++--CAUU-GU++-GU-GGCUUCUUU--+U-A Astereae C---GUU-+GCGGUG-UGCU--CAUG-G+G+-GU-GGC+UCUUU--+U-A Gnaphalieae C---GUUUU+CGGUG-++CU--CAUG-AAAC-U+-UAC+UCUUU--GU-A Inuleae C---GUU-CGCGGUG-UGUG--CAUG-GGA+-G+-GGCUUCUUU--+U-A Senecioneae +---GUU-++CGGUG-AUUG--++++-++AU-CU-+GCUUCUUU--A+-A Anthemideae C---GUU-CGCGGUG-UGCU--CAUG-G+A+-GU-GGCUUCUUU--AU-A Cardueae C---GUU-++CGGUG-UGCA--CA+G-GGUC-GU-GGCCUCUCA--UU-A Vernonieae +++-+UU-+++++++-++++--++++-G+++-+U-++++++UU+--++-A Lactuceae C---GUU+++CGGUG-UGC+--U+++-G+++-GU-+GCCU+CUU--++-A Mutisieae C---G+U-+GCGGUG-UGC+--+G+G-G++++GU-+GCUUCUU---A+-A Barnades. C---GUU-+GCGGUG-++++--+++G-+++C+G+-+GCUUCUU+--++-A 5.8S --> Heliantheae A-+C-+||++++++++++++++++++++++++++++++++++++++++++ Tageteae A-UC-A||+AAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU Eupatorieae A-UC-+||U+++++++++++++++++++++++++++++++++++++++++ Helenieae A-UC-+||UAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU Astereae A-UC-A||CAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU Gnaphalieae A-UC-A||CAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU Inuleae A-CC-A||++++++++++++++++++++++++++++++++++++++++++ Senecioneae A-U+-A||++++++++++++++++++++++++++++++++++++++++++ Anthemideae A-++-+||+++++++++++++++++++++++++++++++++--------- Cardueae A-CC-A||+A+++++++--------------------------------- Vernonieae A-++-A||++++++++++++++++++++++++++++++++++++++++++ Lactuceae A-++-A||C+++++++++++++++++++++++++++++++++++++++++ Mutisieae A-UC-A||++++++++++++++++++++++++++++++++++++++++++ Barnades. A-+C-A||CAAACGACUCUCGGCAACGGAUAUCUCGGCUCACGCAUCGAU Heliantheae ++++++++++++++++++++++++++++++++++++++++++++++++++ Tageteae GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA Eupatorieae ++++++++++++++++++++++++++++++++++++++++++++++++++ Helenieae GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA Astereae GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA Gnaphalieae GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA Inuleae ++++++++++++++++++++++++++++++++++++++++++++++++++ Senecioneae ++++++++++++-+++++++++++++++++++++++++++++++++++++ Anthemideae -----------------+++++++++++++++++++++++++++++++++ Cardueae -------------------------------------------------- Vernonieae ++++++++++++++++++++++++++++++++++++++++++++++++++ Lactuceae ++++++++++++++++++++++++++++++++++++++++++++++++++ Mutisieae ++++++++++++++++++++++++++++++++++++++++++++++++++ Barnades. GAAGAACGUAGCAAAAUGCGAUACUUGGUGUGAAUUGCAGAAUCCCGUGA Heliantheae ++++++++++++++++++++++++++++++++++++++++++++++++++ Tageteae ACCAUCGAGUUUUUGAACGCAAGUUGCGCCCGAAGCCAU++GGUUGAGGG Eupatorieae ++++++++++++++++++++++++++++++++++++++++++++++++++ Helenieae ACCAUCGAGUUUUUGAACGCAAGUUGCGCC+GAAGCC+UCUGG++GAGGG Astereae ACCAUCGAGUUUUUGAACGCAAGUUGCGCCCGAAGCCAUUCGGC+GAGGG Gnaphalieae ACCAUCGAGUUUUUGAACGCAAGUUGCGCC+GAAGCCAUUUGGUUGAGGG Inuleae ++++++++++++++++++++++++++++++++++++++++++++++++++ Senecioneae ++++++++++++++++++++++++++++++++++++++++++++++++++ Anthemideae +++++++++++++++++++++++++++++++++++++++++++++++-++ Cardueae -------------------------------------------------- Vernonieae ++++++++++++++++++++++++++++++++++++++++++++++++++ Lactuceae ++++++++++++++++++++++++++++++++++++++++++++++++++ Mutisieae ++++++++++++++++++++++++++++++++++++++++++++++++++ Barnades. ACCAUCGAGUUUUUGAACGCAAGUUGCGCCC+AAGCCA++CGGC+GAGGG ITS2 --> <-- Helix 2A Heliantheae ++++++++++++++++++++++||+-UCA+G--UCGC-CCC---+A-CCA Tageteae CACGUCUGCCUGGGCGUCACGC||A-UC+UG--U++C-CC+---+A-CCA Eupatorieae ++++++++++++++++++++++||A-UCA+G--U+GC-CCA---CA-+CA Helenieae CACGUCUGCCUGGGCGUCACGC||A-UCACG--UCGC-CC+---CA-C+A Astereae CACGUCUGCCUGGGCGUCACGC||A-UCGCG--UCGC-UCC---C+-C+A Gnaphalieae CACGUCUGCCUGGGCGUCAC+C||A-UCGUG--UCGC-CCC+--+A-CCA Inuleae ++++++++++++++++++++++||+-+++++--UCGC-UCCU--C+-+C+ Senecioneae ++++++++++++++++++++++||+-U+G+G--UC+C-C++---C+-+++ Anthemideae ++++++++++++++++++++++||A-UCGC+--UCGC-CCCC--++-CAA Cardueae ----------------------||A-UCGCG--UCGC-CCC+--+A-+CA Vernonieae ++++++++++++++++++++++||A-U+GC+--UCGC-C+C+--+U-+++ Lactuceae ++++++++++++++++++++++||A-UC+C+--UCGC-CCC+--++-+CA Mutisieae ++++++++++++++++++++++||A-UCGC+--UCGC-CCCC--+A-CAA Barnades. CACG+CUGCCUGGGCGUCACGC||A-UCGCG--UCG+-CCC++-+A-CAA Helix 2A --> Heliantheae +--+++U----CC+---+U+++++G+++U+-----UU+++U-+GGG-GCG Tageteae +--+++U----CC+---++++++GG+U+U+-----+UG+++-+GGG-G+G Eupatorieae +--ACAU----CC+---+++++++++++UG-----+UGUAU-GUGG-GCG Helenieae +--+C+U----CC+---++++++GGAU+++-----+U+G+U-G+GG-GCG Astereae +--++++----+C+---UU--++GG-AUG+-----UUGG+U-GGG+-GCG Gnaphalieae +--+++U----C++---UUA-+++GAUGUU-----U+GUG+-+GGG-GCG Inuleae U--GC+U----CCU---+-AAA+GGGUG+G-----C++GAU-A+GA-GCG Senecioneae +-++C++--++++U---U+A+GGGG+UGU+-----U++++U-GGGG+G+G Anthemideae +--U+++-----++---+++++++++++++-----U+U+U+-GGGG-GCG Cardueae +--++++----C++---++++G++++++GU-----+U+GU+-UGGG-+CG Vernonieae +--GC+U--+++++---A+++++A++C++------+U++UU-GGGG-GCG Lactuceae +--+C+U+---C++---+++++G+++++++-----+G+++U-+GGG-GCG Mutisieae +--G++U+---CC+---++++GG+++++++++---+++G++-GGGG-GCG Barnades. C--+CA++---C++---+++++GCG++++------++++++-+GGG-GCG <-- Helix 2B --> Heliantheae G-A+--AU-U-GGUC-UC-C+G-UGCC-+AU---GG+G-+GG-UU-GGCC Tageteae +-A+--AU-U-GGUC-UC-C+G-UGCC-+++++-GG+G-++G-UU-GGCC Eupatorieae G-AG--AC-U-GGUC-UC-CUG-UGCC-+A+---GGUG-CGG-UU-GGCC Helenieae G-AG--AU-U-GGUC-UC-C+G-UG++-++U---G+++-+GG-UU-GGCC Astereae G-AU--A+-U-GGUC-UC-CCG-UUU+UCAC---CGAG-CGG-UU-GGCC Gnaphalieae G-AG--AU-U-GGUC-UC-CCG-+UUC-U++---GAUA-CGG-UU-GGCC Inuleae G-AU--A+-U-GGUC-UC-CCG-UGCC-UA+---GGUG-+GG-UU-GGC+ Senecioneae G-AG--AU-U-GGU+-UC-CCG-UU+C-U++---G+UG-CGG-UU-GGCU Anthemideae G-AU--AU-U-GGUC-UC-CCG-UGCU-CA+---GG+G-UGG-UU-GGCC Cardueae G-A+--A+-U-GGUC-UC-CCG-UG++-+++---GG+G-+GG-UU-GGCC Vernonieae G-A+--AU-U-GG+C-UC-CCA-UGC+-+A+---GG+G-UGG-UU-GGCC Lactuceae G-A+--AU-U-GG+C-UC-CCG-U+C+-U++---+GU+-+GG-UU-GGCC Mutisieae G-A+--A+-U-GG+C-U+-CCG-UGC+-++++--GG+G-++G-UU-GGCC Barnades. G-A+--++-U-GG+C-UC-CCG-UG+C-+++++-+G+G-+GG-+U-GGCC <-- Helix 2C Heliantheae UA-AA-UA+-+AGUC+CCU-C++-GA------G+-GAC-GCACG+-C-U- Tageteae UA-AA++A+-GAGUC++C+-+AA+++------++-GA+-GCACGA-C-U- Eupatorieae CA-AA-+A+-GAG+CC++U-UAA-GA++----G+-GAC-GCACGA-C-U- Helenieae UA-AA-UA+-GAGUC+++U-C+A-GA------+G-GA+-GCACGA-C-U- Astereae +A-AA-+AA-+AG++CC++-UU--GA------CG-GG+-+CA+GA-C-U- Gnaphalieae +A-AA-UAG-GAGUCCCCA-UC--GA------UG-GAC-GCACGA-C-U- Inuleae AA-AA-UAG-+AGUCU+CU-U+--GA------+G-GAC-ACACG+-C-A- Senecioneae +A-A+-U++-GA+U+C++U-U++++A------+G-+AC-+C+CGA-U-U- Anthemideae +A-AA-UA+-GAGU-CCCU-UC--GA------UG-GAC-GCAC+AAC-U- Cardueae UA-AA-AA+-GAGUCCCCU-U+--G+------+G-G++-GCACGG-C-U- Vernonieae +A-AA-++G-AAGUCCU++-UC--G+------U+-G+U-GCA+GA-C-U- Lactuceae +A-AA-+++-GAGU++CCU-++--GG------+G-GA+-GCAC+A-C-U- Mutisieae UA-AA-+++-GAGUCC++U-U+--G+------+G-GAC-GCAC++-C-U- Barnades. +A-AA++++-G+GUC+C++-++--GA------++-+++-+CAC++-C-+- Helix 2C --> Heliantheae ---AGUGGUGGUU-GAUAA-GAC+---G-UCGU+UC--G+GU-+GUG+GU Tageteae ---AGUGGUGGUU-UGUAA-+A+A---G-UCGUCUC--GUGU-UGUG++U Eupatorieae ---GGUGGUGGUU-GAUUA-+AC+---G-UCGUCUC--GUGU-CGUG+G+ Helenieae ---AGUGGUGGUU-GAU+A-+ACA---G-UCGUCUC--GUGU-CGUG+GU Astereae ---A+UG+UGGU+-GACAA-AACC-----++GAAU+--++GU-+G+GUGU Gnaphalieae ---AGUGGUGGUU-GACAA-AACC---U-UCGUCUU--G+GU-+GUGCGU Inuleae ---AGUGGUGGUU-GA+AA-AACC---U++AGUCUC--GUGU-+GUGUGU Senecioneae ---AGUGGUGGUU-G++AA-GACC---+-UCU+AU+--GAGU-+GUG+GU Anthemideae ---AGUGGUGGUC-GUAAA-AACC---C-UCGU++U--UUGU-U++G+G+ Cardueae ---AGUGGUGGUU-GU+AA-GGCC---U-UCGUAUC--GAGC-CGUG+-- Vernonieae ---A+UGGUGGUU-+++AA-GACC---U-++G+U++--++GU-+++GU++ Lactuceae ---AGUGGUGGUU-GAA+A-GACC---C-UCGUCUU--+UGU-+GUG+GU Mutisieae ---AGUGGUGGUU-G+CAA+G+CC---+-U+++UUC--+UG+-CGUGCGU Barnades. ---AGUGGUGG++-+++AA-+GCC-----++AGUUC--GUGU-CGUG++U Heliantheae U-+---+++--+++U+++G++++++++CU+--+U+++-++A--CC+-+++ Tageteae +-+---U+A+-+++++++++G++A++++U+--U+A++-+++++CCC--++ Eupatorieae +-U---+GA--UUCUUG-A++G+A+A+++++-UU++A-+UA--CCC-U+A Helenieae U-+---++A--+UCUUGA+G+++A++ACUC--UUAAA-++A--CCC-+G+ Astereae C-U---+G---UCAAAA-GG+UGC+UCUUA--++----+GA--CCC--AA Gnaphalieae +-+---U+AG-UCGUA+-GGGAAGAU-CUC--UUUAA-AGA--CCC-CAA Inuleae C-C---+GA+-UUGUAA-GCGAAGA-CC+++-UA++--+UA--CCC-UAA Senecioneae U-C---++A+-++G+AA--+G+A+A+-C+C--U++AA-U+A--+CC-U+A Anthemideae +-+---+++G-UCG+AA-GG+AA++--CUC--U+++A-A+A--CCC-+AA Cardueae ------++A+-++G++A-GGGA+++G-CUC--U++AA-AGA--+CC-+AA Vernonieae C-+---++AG-+CG+AA-G++A+A++-+UU+-+++AA-AG+--CC+-U++ Lactuceae +-G---U+AG-C+G++A-+GGA+G++-+U+--+++++-AGA--CCC-+A+ Mutisieae +-+---+++G-UCGC+A-++++++-G++++++G++A--AGA--CCC-+AA Barnades. +-+---++++-+CG+++--+GG++++-CUC+-++++--AGA--CCC+++A <-- Helix 2D --> Heliantheae +--G+-GU-UGU-C-+U-++-GA+GA+GC-UUC-GA++|| Tageteae U--G+-+U-UGU-C-UU-++-GA++AUGC-UUC-GAUC|| Eupatorieae U--G+-G+-+GU-C-UU-+U-G++++CUC-UU+-GAU+|| Helenieae +--G+-GU-UGU-C-UU-+U-GAUG++GC-UU+-GA--|| Astereae +--GC-GU-UGU-+-+U-++-+A+GA+GC-UUC-GA++|| Gnaphalieae U--G+-GU-UGU-C-UU-U+-GAUGAC+C-UUC-G+++|| Inuleae C--GU-GU-+GU-C-++-+U-GAC+ACGC-UUC-GACC|| Senecioneae U--GU-G+-C+U-+-U+-+U-+A+GA+GC-U++-GA++|| Anthemideae +--GU-GU-+GU-C-UU-++-GA+GACGC-UUC-GA++|| Cardueae +--GU-GU-+GU-C-UU-++-GACGA+GC-UUC-GA--|| Vernonieae U--G+-+U-+G+-+-U+-++-++++A+G+-UUC-G+++|| Lactuceae U--GU-+U-+GU-+-++-++-+A+GAU++-+UC-GAC+|| Mutisieae +--G+-GU-CGU-C-++-++-GACG++G+-UU+-GACC|| Barnades. +--++-G+-+++-C-U++++-G++G+++C-U+C-GAC+||